
Á. L. Viloria
82
It was Trebbau’s well known inclination to make zoology 
available to ordinary people. Accordingly, he took the ini-
tiative to deeply update the scientic content of the book, 
incorporating an appreciable amount of novel informa-
tion, recent or previously omitted, that has been revealed 
through eld experiences and scientic studies published 
during the last forty years. For this task, he requested the 
collaboration of four Venezuelan herpetologists, Tito R. 
Barros, Omar E. Hernández, Hedelvy J. Guada and Gil-
son A. Rivas, who with their recognized experience in 
nature surveying and their good knowledge of turtles, 
were kind enough to help worthily to one of the masters 
of eld vertebrate zoology of Venezuela. Trebbau also 
thought of adding an original appendix that consists of a 
review on turtles in the fossil record of Venezuela, written 
by paleontologists Jorge D. Carrillo Briceño and Marcelo 
R. Sánchez Villagra. All chapters, in addition to being il-
lustrated with the beautiful paintings made by Giorgio 
Voltolina for the plates of the rst edition, include a num-
ber of good photographic images provided by more than 
30 contributors. Standing out for their imaginative style, 
there are two precious pictorial reconstructions of fossil 
taxa in their palaeoenvironments, illustrating the paleon-
tological appendix. ese were produced by the biologi-
cal artist Jorge A. González (pp. 215, 216)
e main involvement of Peter Pritchard (1943-2020) 
in this editorial project dates back to the 1970s. His career 
as a world expert on turtles and his friendly initiative pro-
vided much more enthusiasm for Trebbau to advance and 
complete this work. I personally remember the anecdote in 
which both authors discovered the peculiar western Ven-
ezuelan endemic species, Mesoclemmys zuliae, while car-
rying out a routine to take morphometric measurements 
of captive chelonians in the main pond of the Zoológico 
Parque Sur de Maracaibo (the zoo of the second largest 
city in Venezuela). Before his astonished eyes a specimen 
of this unknown toad-headed turtle appeared. ey im-
mediately initiated a guided search, which led them to the 
discovery of its natural habitat in the swamps and streams 
of the neighborhood of El Guayabo, in the south of Lake 
Maracaibo region. Listening to their story produced the 
greatest possible enthusiasm in younger zoology amateurs 
because of the evident possibilities for discovery still of-
fered by some underexplored tropical regions.
is book begins with a heartfelt dedication to the 
memory of Saúl Gutiérrez (1960-2012), a respected col-
league dedicated to the science of zoological parks, and 
a disciple of the rst author. A presentation written by 
Carlos Rivero Blanco brings historical memories with an 
inevitable tone of admiration to the authors, of whom he 
was a close collaborator for several years. Vivian P. Páez 
(University of Antioquia, Colombia) makes a precise 
summary of the qualities of the work in a foreword that 
covers virtually all what is needed to say. It is in fact a con-
cise book review.
Expanding on Páez’s opinion in relation to the content 
of the book, I consider that it introduces the reader very 
well to the general knowledge of turtles as living beings 
and to their biology. Later on, it focuses on the physical 
characteristics of their morphology and anatomy, empha-
sizing graphically the anatomical nomenclature to describe 
the carapace plates, which forms the basis of the descrip-
tive terminology used throughout the work. A complete 
count of the 22 living species of turtles (terrestrial, fresh-
water and marine) known in the territory of Venezuela is 
presented, they are included in een genera and eight 
families. Each taxon is described and illustrated individu-
ally, with input of data on its geographical distribution 
- including a map for each taxon in what corresponds to 
Venezuela -, its habitat, its diet and its reproductive biol-
ogy. It is also widely discussed what is known about the 
exploitation of each species by human populations, for 
consumption as a source of protein or as a supplier of raw 
materials for ornaments such as hawksbill, nowadays fallen 
into disuse due to the supervening threats because of the 
slaughter for these purposes. is section leads directly to 
long considerations on the conservation status of each tax-
on, a subject that the authors handle with great erudition, 
since they were pioneers on this eld.
Towards the end of the book there are a couple of brief 
but important sections: the rst mentions the species of 
turtles not yet detected, but whose presence is predictable 
in some part of the territory of Venezuela, as could be in-
ferred from their general distribution; and in the second, 
explanations are presented on the erroneously cited spe-
cies for the country.
As expected from a revised and updated edition, the 
list of bibliographic references at the end is complete and 
exhaustive. It takes time to build up such a collection of 
sources. e reader is advised to pay attention to the many 
little-known papers and monographs in this section.
Finally, a small but very pleasant additional detail refers 
to an unexpected cultural element. It is the representation 
of native turtles in philately and notaphily in Venezuela. 
e book contains reproductions of images of Venezuelan 
postal stamps and banknotes, whose main motifs are the 
national emblematic turtles. ese are past eorts that 
have been made to spread popular knowledge of the au-
tochthonous cheloniofauna of Venezuela and to promote 
its conservation worldwide.